Reference
Osvaldo A. Reig, A. Perez-Zapata, Marisol Aguilera, A. Ferrer (1985): Cytogeneics and karyosystematics of South American oryzomyine rodents (Cricetidae: Sigmodontinae)
I. A species of Oryzomys with a low chromosome numer from Northern Venezuela – Mammalian Biology (früher Zeitschrift für Säugetierkunde) – 51: 368 - 378.