Reference
Fernando Leal, Camila Zornosa-Torres, Guilherme Augusto-Alves, Simone Dena, Tiago Leite Pezzuti, Felipe Leite, Luciana Bolsoni Lourenco, Paulo Garcia, Luis Felipe Toledo (2021): Head in the clouds: a new dwarf frog species of the Physalaemus signifer clade (Leptodactylidae, Leiuperinae) from the top of the Brazilian Atlantic Forest – European Journal of Taxonomy – 0764: 119 - 151.