Reference
Lucindo Gonzales, Stefan Lötters, Steffen Reichle (1999): On the dendrobatid frogs from Bolivia: rediscovery of Epipedobates bolivianus (Boulenger, 1902), first record of Colostethus brunneus (Cope, 1887) and comments on other species (Anura: Dendrobatidae). – Herpetozoa – 12_3_4: 179 - 186.