Reference
Oleg N. Kabakow (1998): Studies on the Coprophagous Scarab Beetles (Coleptera, Scarabaeidae) from Vietnam in the collections of Zoological Institute St. Petersburg and of the Naturkundemuseum Erfurt – Veröffentlichungen des Naturkundemuseums Erfurt (in Folge VERNATE) – 17: 39 - 58.