Reference

Holger Dombrow (2001): Revision of the Genus Gymnoloma Burmeister 1844 with observations on three new genera (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). 14th Contribution to the knowledge of African Hopliini – Coleoptera - Schwanfelder Coleopterologische Mitteilungen – 5: 107 - 148.