Reference

Paolo Neri, Luca Toledano (2018): Notes on the genus Bembidion Latreille, 1802, subgenus Ocyturanes Müller-Motzfeld, 1986, with particular reference to the "signatipenne group" and description of a new species from Iran (Coleoptera, Carabidae, Bembidiini) – Entomologische Blätter – 114: 301 - 308.