Reference
Zachary H. Griebenow, Adrian Richter, Thomas van de Kamp, Evan P. Economo, Ziv E. Lieberman (2023): Comparative morphology of male genital skeletomusculature in the Leptanillinae (Hymenoptera: Formicidae), with a standardized muscular terminology for the male genitalia of Hymenoptera – Arthropod Systematics and Phylogeny – 81: 945 - 1018.