Reference

C.J. Wilson, Robert B. Angus (2004): A chromosomal analysis of ten European species of Aphodius ILLIGER, subgenera Acrossus MULSANT, Nimbus MULSANT & REY and Chilothorax MOTSCHULSKY (Coleoptera: Aphodiidae). – Koleopterologische Rundschau – 74_2004: 367 - 374.