Reference
C.J. Wilson, Robert B. Angus (2005): A chromosomal analysis of four European species of Aphodius ILLIGER, subgenera Colobopterus MULSANT, Coprimorphus MULSANT, Otophorus MULSANT and Teuchestes MULSANT (Coleoptera: Aphodiidae). – Koleopterologische Rundschau – 75_2005: 319 - 323.